† denotes equal contribution.



  1. Reconceptualizing beta diversity: a hypervolume geometric approach
    Chuliang Song†, Muyang Lu†, Joseph R. Bennett, Benjamin Gilbert, Marie-Josée Fortin, and Andrew Gonzalez
    submitted, 2024
  2. Landscape ecological consolidation intensifies zoonotic host diversity loss and disease transmission in human habitats
    Shan Pei, Pengbo Yu, Jayna Raghwani, Yuxin Wang, Ziyan Liu, Yidan Li, and 17 more authors
    submitted, 2024
  3. Soil microbial influences over coexistence potential in multispecies plant communities in a subtropical forest
    Hangyu Wu, Weitao Wang, Tingting Wu, Zijing Luo, Wei Lin, Hanlun Liu, and 7 more authors
    submitted, 2024



  1. Trophic tug-of-war: Coexistence mechanisms within and across trophic levels
    Chuliang Song†, and Jurg W. Spaak†
    Ecology Letters, 2024


  1. Mean species responses predict effects of environmental change on coexistence
    Frederik De Laender, Camille Carpentier, Timoteo Carletti, Chuliang Song, Samantha L. Rumschlag, Michael B. Mahon, and 3 more authors
    Ecology Letters, 2023
  2. Rapid monitoring of ecological persistence
    Chuliang Song†, Benno I. Simmons†, Marie-Josée Fortin, Andrew Gonzalez, Christopher N. Kaiser-Bunbury, and Serguei Saavedra
    Proceedings of the National Academy of Sciences, 2023


  1. Generalism drives abundance: A computational causal discovery approach
    Chuliang Song, Benno I. Simmons, Marie-Josée Fortin, and Andrew Gonzalez
    PLOS Computational Biology, 2022
  2. Metapopulation persistence can be inferred from incomplete surveys
    Chuliang Song, Marie-Josée Fortin, and Andrew Gonzalez
    Proceedings of the Royal Society B, 2022
  3. Synthesizing the effects of individual-level variation on coexistence
    Simon Maccracken Stump, Chuliang Song, Serguei Saavedra, Jonathan M Levine, and David A Vasseur
    Ecological Monographs, 2022


  1. Coexistence holes characterize the assembly and disassembly of multispecies systems
    Marco Tulio Angulo, Aaron Kelley, Luis Montejano, Chuliang Song†, and Serguei Saavedra†
    Nature Ecology & Evolution, 2021
  2. Bridging parametric and nonparametric measures of species interactions unveils new insights of non-equilibrium dynamics
    Chuliang Song, and Serguei Saavedra
    Oikos, 2021
  3. Understanding the emergence of contingent and deterministic exclusion in multispecies communities
    Chuliang Song, Lawrence H Uricchio, Erin A Mordecai, and Serguei Saavedra
    Ecology Letters, 2021
  4. Untangling the complexity of priority effects in multispecies communities
    Chuliang Song, Tadashi Fukami, and Serguei Saavedra
    Ecology Letters, 2021
  5. Merging dynamical and structural indicators to measure resilience in multispecies systems
    Lucas P Medeiros†, Chuliang Song†, and Serguei Saavedra
    Journal of Animal Ecology, 2021


  1. Disentangling the effects of external perturbations on coexistence and priority effects
    Chuliang Song, Rudolf P. Rohr, David Vasseur, and Serguei Saavedra
    Journal of Ecology, 2020
  2. Telling ecological networks apart by their structure: An environment-dependent approach
    Chuliang Song, and Serguei Saavedra
    PLOS Computational Biology, 2020
  3. Structural stability: concepts, methods, and applications (in Chinese)
    Chuliang Song
    Biodiversity Science, 2020
  4. Towards a probabilistic understanding about the context-dependency of species interactions
    Chuliang Song, Sarah Von Ahn, Rudolf P Rohr, and Serguei Saavedra
    Trends in Ecology and Evolution, 2020
  5. Accelerating The Emergence Of Order In Swarming Systems
    Yandong Xiao, Chuliang Song, Liang Tian, and Yang-Yu Liu
    Advances in Complex Systems, 2020


  1. On the consequences of the interdependence of stabilizing and equalizing mechanisms
    Chuliang Song†, György Barabás†, and Serguei Saavedra
    The American Naturalist, 2019
  2. Beware z-scores
    Chuliang Song, Rudolf P Rohr, and Serguei Saavedra
    Journal of Animal Ecology, 2019


  1. Rethinking the importance of the structure of ecological networks under an environment-dependent framework
    Simone Cenci†, Chuliang Song†, and Serguei Saavedra
    Ecology and evolution, 2018
  2. Will a small randomly assembled community be feasible and stable?
    Chuliang Song, and Serguei Saavedra
    Ecology, 2018
  3. Structural changes within trophic levels are constrained by within-family assembly rules at lower trophic levels
    Chuliang Song, Florian Altermatt, Ian Pearse, and Serguei Saavedra
    Ecology Letters, 2018
  4. A guideline to study the feasibility domain of multi-trophic and changing ecological communities
    Chuliang Song, Rudolf P Rohr, and Serguei Saavedra
    Journal of Theoretical Biology, 2018
  5. Structural stability as a consistent predictor of phenological events
    Chuliang Song, and Serguei Saavedra
    Proceedings of the Royal Society B: Biological Sciences, 2018


  1. Why are some plant–pollinator networks more nested than others?
    Chuliang Song, Rudolf P Rohr, and Serguei Saavedra
    Journal of Animal Ecology, 2017


  1. Existence of positive solutions for an approximation of stationary mean-field games
    Nojood Almayouf, Elena Bachini, Andreia Chapouto, Rita Ferreira, Diogo Gomes, Daniela Jordão, and 19 more authors
    Involve, a Journal of Mathematics, 2016